CASSIDINAE - TORTOISE BEETLES
Named for their distinctive defensive behaviour of drawing in the head and legs under the armoured elytra and prothorax, the Tortoise Beetles are for the most part instantly recognisable. Ths Chrysomelid subfamily Hispinae is no longer recognised and is now considered a tribe of the Cassidinae.
Online Resources
Cassidinae of the World: An Interactive Manual Professor Lech Borowiec´s web page this is highly recommended for anybody with an interest in this group - thanks to Professor Borowiec the best studied of the Chrysomelid subfamilies - featuring species lists, identification keys and images of a large number of specimens.
Click on the images to enlarge them.
Thanks to Professor Lech Borowiec for his assistance with identification of some of the images in this gallery.
Designed by Paul Smith 2006. This website is copyrighted by law.
Material contained herewith may not be used without the prior written permission of FAUNA Paraguay.
Photographs on this page were taken by Paul Smith, Pier Cacciali and Alberto Esquivel and are used with their permission.
![FIGURE 1 FIGURE 1](Botonochara decemverrucata Cassidinae san raf oct 08.jpg) | FIGURE 1 |
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![FIGURE 2 FIGURE 2](Charidotella sp chrysomelidae san rafael 16 nov 07.jpg) | FIGURE 2 |
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![FIGURE 3 FIGURE 3](Coptocycla adamantina Cassidinae 7 nov 08 tirol 2.jpg) | FIGURE 3 |
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![FIGURE 4 FIGURE 4](Coptocyla (Psalidonota) contemta pier cacciali 9 june 08 proc.jpg) | FIGURE 4 |
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![FIGURE 5 FIGURE 5](Corialomela brunnea Thunberg 1821 chrysomelidae BAHIA NEGRA sep 06.jpg) | FIGURE 5 |
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![FIGURE 6 FIGURE 6](Corialomela thoracica Cassidinae beetle san raf 21 mar 08 david gill.jpg) | FIGURE 6 |
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![FIGURE 7 FIGURE 7](Paraselenis flava Cassidinae san raf oct 08.jpg) | FIGURE 7 |
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![FIGURE 8 FIGURE 8](Stolas festiva Klug 1829 chrysomelidae san raf march 07.jpg) | FIGURE 8 |
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![FIGURE 9 FIGURE 9](Storas lacordairei prob chrysomelidae 2 san rafael 16 nov 07.jpg) | FIGURE 9 |
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![FIGURE 10 FIGURE 10](Stolas redtenbacheri cassidinae chrysomelid aem proc 8 nov 07.jpg) | FIGURE 10 |
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![FIGURE 11 FIGURE 11](Stolas redtenbacheri cassidinae chrysomelid AEM proc 16 nov 07.jpg) | FIGURE 11 |
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![FIGURE 12 FIGURE 12](Cassidinae - Stolaini larva san rafael 16 nov 07.jpg) | FIGURE 12 |
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FIGURE 1 - Botonochara decemverrucata - PROCOSARA, PN San Rafael (Carlos Aguilar Julio October 2008 - ECOSARA Biodiversity Database).
FIGURE 2 - Charidotella sp. (in C.vinula species group) - PROCOSARA, PN San Rafael (Paul Smith November 2007 - ECOSARA Biodiversity Database).
FIGURE 3 - Coptocyla adamantina - Hotel Tirol, Departamento Itapúa (Paul Smith November 2008).
FIGURE 4 - Coptocyla (Psalidonota) contemta - PROCOSARA, PN San Rafael (Pier Cacciali June 2007 - ECOSARA Biodiversity Database).
FIGURE 5 - Coraliomela brunnea - Bahía Negra, Departamento Alto Paraguay (Paul Smith September 2006).
FIGURE 6 - Coraliomela thoracia - PROCOSARA, PN San Rafael (David Gill March 2008 - ECOSARA Biodiversity Database).
FIGURE 7 - Paraselenis flava - PROCOSARA, PN San Rafael (Carlos Aguilar Julio October 2008 - ECOSARA Biodiversity Database).
FIGURE 8 - Stolas festiva - PROCOSARA, PN San Rafael (Paul Smith March 2007 - ECOSARA Biodiversity Database).
FIGURE 9 - Stolas sp. probably lacordairei blue form (Cassidinae) - PROCOSARA, PN San Rafael (Paul Smith November 2007 - ECOSARA Biodiversity Database).
FIGURE 10 - Stolas redtenbacheri - PROCOSARA, PN San Rafael (Alberto Esquivel November 2007 - ECOSARA Biodiversity Database).
FIGURE 11 - Stolas redtenbacheri - PROCOSARA, PN San Rafael (Alberto Esquivel November 2007 - ECOSARA Biodiversity Database).
FIGURE 12 - Larva of Tortoise Bug (Tribe Stolaini) - PROCOSARA, PN San Rafael (Paul Smith November 2007 - ECOSARA Biodiversity Database).