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Material contained herewith may not be used without the prior written permission of FAUNA Paraguay.
Photographs on this page were taken by José Luis Cartes and are used with permission.
SOUTHEASTERN FOUR-EYED OPOSSUM Philander frenatus
Smith P 2009 - FAUNA Paraguay Handbook of the Mammals of Paraguay Number 9 Southeastern Foure-eyed Opossum Philander frenatus
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Opossums are the American branch of the marsupial order - they raise their young in a pouch. This is a rarely encountered species in Paraguay due to its nocturnal habits and semi-arboreal nature. They are excellent swimmers. The common name refers to the four white spots above the eyes. The species has been recently split from the more widespread Grey Four-eyed Opossum Philander opossum.
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FIGURE 1 - (FPMAM12PH) Juvenile, location unknown (José Luis Cartes undated).
FIGURE 2 - (FPMAM13PH) Juveniles from same litter (José Luis Cartes undated).
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LINKS TO DOWNLOADABLE OPEN ACCESS REFERENCES USED IN THE PREPARATION OF THE FAUNA PARAGUAY HANDBOOK OF THE MAMMALS OF PARAGUAY SPECIES ACCOUNT FOR THIS SPECIES:
Allen JA 1900 - Note on the Generic Names Didelphis and Philander - Bulletin AMNH 13: p185-190.
Brown BE 2004 - Atlas of New World Marsupials - Fieldiana Zoology 102.
Burmeister H 1854 Systematische Uebersicht der Thiere Brasiliens: Welche während einer Reise durch die Provinzen von Rio de Janeiro und Minas Geraës Gesammlt oder beobachtet wurden Vol 1 - G.Reimer, Berlin.
Cáceres NC 2004 - Diet of Three Didelphid Marsupials (Mammalia: Didelphimorphia) in Southern Brazil - Mammalian Biology 69: p430-433.
Carvalho FMV de, Delciellos AC, Vieira MV 2000 - Medidas Externas dos Miembros de Marsupiais Didelfidios: Uma Comparação com Medidas do Esqueleto Pós-Craniano - Boletim do Museu Nacional, Rio de Janeiro 438.
Carvalho FMV de, Pinheiro PS, Fernández FAS, Nessimian JL 1999 - Diet of Small Mammals in Atlantic Forest in Southeastern Brazil - Revista Brasileira de Zoociencias 1: p91-101.
Ceotto P, Finotti F, Santori R, Cerqueira R 2009 - Diet Variation of the Marsupials Didelphis aurita and Philander frenatus (Didelphimorphia: Didelphidae) in a Rural Area of Rio de Janeiro State, Brazil - Mastozoologia Neotropical 16.
Cunha AA, Vieira MV 2002 - Support Diameter, Incline and Vertical Movements of Four Didelphid Marsupials in the Atlantic Forest of Brazil - Journal of Zoological Society of London 258: p419-426.
Delciellos AC, Vieira MV 2006 - Arboreal Walking Performance in Seven Didelphid Marsupials as an Aspect of Their Fundamental Niche - Austral Ecology 31: p449-457.
Delciellos AC, Vieira MV 2007 - Stride Lengths and Frequencies of Arboreal Walking in Seven Species of Didelphid Marsupials - Acta Theriologica 52: p101-111.
Delciellos AC, Vieira MV 2009 - Allometric, Phylogenetic and Adaptive Components of Climbing Performance in Seven Species of Didelphid Marsupials - Journal of Mammalogy 90: p104-113.
Hershkovitz P 1997 - Composition of the Family Didelphidae Gray 1821 (Didelphoidea: Marsupialia) with a Review of the Morphology and Behaviour of the Included Four-eyed Pouched Opossums of the Genus Philander Tiedemann 1808 - Fieldiana 86.
Moura MC, Capparelli AC, Freitas SR, Vieira MV 2005 - Scale Dependent Habitat Selection in Three Didelphid Marsupials Using the Spool and Line Technique in the Atlantic Forest of Brazil - Journal of Tropical Ecology 21: p337-342.
Myers P, Espinosa R, Parr CS, Jones T, Hammond GS, Dewey A 2006 - The Animal Diversity Web (online). Accessed December 2007.
Vieira MV 1997 - Body Size and Form in Two Neotropical Marsupials, Didelphis aurita and Philander opossum (Marsupialia: Didelphidae) - Mammalia 61: p245-254.
Vieira MV, Cunha AdeA 2008 - Scaling Body Mass and Use of Space in Three Species of Marsupials in the Atlantic Forest of Brazil - Austral Ecology 33: p872-879.
