PYRGINAE Flat-winged Skippers FAUNA PARAGUAY

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Gallery 1 Achylodes, Anastrus, Anisochoria and Antigonus
FIGURE 1a - Giant Sicklewing Achlyodes busirus - Encarnación, Departamento Itapúa (Paul Smith June 2007).
FIGURE 1b - Common Sicklewing Achylodes mithridates male - Encarnación, Departamento Itapúa (Paul Smith June 2007).
FIGURE 1c - Common Sicklewing Achylodes mithridates female - PROCOSARA, PN San Rafael (Moriz Stiefel April 2008).
FIGURE 1d - Anastrus semipternus - Encarnación, Departamento Itapúa (Paul Smith May 2009).
FIGURE 1e - Anisochoria pedaliodina extincta - Estancia Kanguery, PN San Rafael (Paul Smith November 2007).
FIGURE 1f - Cut-wing Skipper Anisochoria sublimbata - PROCOSARA, PN San Rafael (Paul Smith March 2007).
FIGURE 1g - Antigonus erosus male - PROCOSARA, PN San Rafael (Paul Smith March 2008).
FIGURE 1h - Antigonus erosus female - PROCOSARA, PN San Rafael (Paul Smith March 2008).
FIGURE 1i - Antigonus nearchus - PROCOSARA, PN San Rafael (Paul Smith November 2007).

Gallery 5 Gesta, Gorgythion, Grais and Helias

FIGURE 5a - Gesta gesta - Estancia Laguna Blanca, Departamento San Pedro (Paul Smith October 2008).
FIGURE 5b - Gorgythion begga - PROCOSARA, PN San Rafael (Jeni Oborn June 2008).
FIGURE 5c - Gorgythion beggina - Encarnación, Departamento Itapúa (Paul Smith April 2008).
FIGURE 5d - Grais stigmaticus - PROCOSARA, PN San Rafael (Laura Tensen March 2008).
FIGURE 5e - Helias phalaenoides - PROCOSARA, PN San Rafael (upper Moriz Stiefel, lower Paul Smith May 2008).

Gallery 10 Sostrata, Staphylus, Timochares, Trina and Typhedanus
FIGURE 10a - Lavender Skipper Sostrata bifasciata - PROCOSARA, PN San Rafael (Paul Smith November 2007).
FIGURE 10b - Sostrata cronion - PROCOSARA, PN San Rafael (Sylvia Qu May 2008).
FIGURE 10c - Staphylus ascalon - PROCOSARA, PN San Rafael (Paul Smith November 2007).
FIGURE 10d - Three-banded Skipper Timochares trifasciata - PROCOSARA, PN San Rafael (upper Paul Smith March 2007 lower Alberto Esquivel December 2007).
FIGURE 10e - Geometric Skipper Trina geometrina - PROCOSARA, PN San Rafael (Paul Smith February 2008).
FIGURE 10f - Typhedanus undulatus - PROCOSARA, PN San Rafael (Paul Smith May 2008).

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Gallery 7 Mylon, Narcosius, Nascus and Oechydrus
FIGURE 7a - Mylon maimon - PROCOSARA, PN San Rafael (Paul Smith upper March 2007; lower November 2007).
FIGURE 7b - Mylon pelopidas - location unknown (Ulf Drechsel www.pybio.org).
FIGURE 7c - Narcosius parisi - Hotel Tirol, Departamento Itapúa (Paul Smith September 2007).
FIGURE 7d - Common Scarlet-Eye Nascus phocus female - Encarnaciòn, Departamento Itapúa (Paul Smith upper October 2006 lower April 2008).
FIGURE 7e - Nisionades macarius - location unknown (Sergio Rios undated).
FIGURE 7f - Oechydrus chersis - Hotel Tirol, Departamento Itapúa (Paul Smith September 2007).

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Gallery 11 Urbanus
FIGURE 11a - Urbanus chalco - location unknwon (Ulf Drechsel www.pybio.org).
FIGURE 11b - Lilac-banded Long-tailed Skipper Urbanus dorantes - PROCOSARA, PN San Rafael (Paul Smith May 2008).
FIGURE 11c - Urbanus doryssus - PROCOSARA, PN San Rafael (Jeni Oborn June 2008).
FIGURE 11d - Urbanus esta - PROCOSARA, PN San Rafael (Moriz Stiefel April 2008).
FIGURE 11e - Urbanus procne - Encarnación, Departamento Itapúa (Paul Smith May 2007).
FIGURE 11f - Urbanus pronta - PROCOSARA, PN San Rafael (Nick Simms February 2008).

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Gallery 2 Astraptes and Autochton
FIGURE 2a - Yellow-tailed Skipper Astraptes anaphus - Encarnación, Departamento Itapúa (Paul Smith May 2009).
FIGURE 2b - Chalk-spotted Blue Skipper Astraptes creteus - Encarnación, Departamento Itapúa (Paul Smith April 2008).
FIGURE 2c - Astraptes elorus - PROCOSARA, PN San Rafael (Paul Smith March 2008).
FIGURE 2d - Astraptes fulgerator - PROCOSARA, PN San Rafael (Laura Tensen April 2008).
FIGURE 2e - Astraptes fulgor - PROCOSARA, PN San Rafael (Paul Smith May 2008).
FIGURE 2f - White-banded Skipper Autochton zarex - upper PROCOSARA, PN San Rafael (Paul Smith March 2008), lower Hotel Tirol, Departamento Itapúa (Paul Smith September 2007).

Gallery 3 Bolla, Bungalotis, Camptopleura, Celaenorrhinus, Chioides, Chiomara, Clito and Codatractus
FIGURE 3a - Bolla sp. - PN Tte Enciso, Departamento Boquerón (Paul Smith October 2008).
FIGURE 3b - Bungalotis astylos female - PROCOSARA, PN San Rafael (Paul Smith September 2008).
FIGURE 3c - Camptopleura auxo - Estancia Laguna Blanca, Departamento San Pedro (Paul Smith February 2010).
FIGURE 3d - Obstinate Skipper Celaenorrhinus similis - PROCOSARA, PN San Rafael (upper Sylvia Qu June 2008; lower Moriz Stiefel May 2008).
FIGURE 3e - White-striped Long-tailed Skipper Chioides catillus - PN Tte Enciso, Departamento Boquerón (Paul Smith July 2006).
FIGURE 3f - White-patched Skipper Chiomara asychis - PN Tte Enciso, Departamento Boquerón (Paul Smith July 2006).
FIGURE 3g - Chiomara mithrax - Estancia Laguna Blanca, Departamento San Pedro (Paul Smith October 2008).
FIGURE 3h - Clito bubulus female - Arroyo Clementina, Departamento San Pedro (Paul Smith October 2008).
FIGURE 3i - Gold-bordered Long-tailed Skipper Codatractus aminias - PROCOSARA, PN San Rafael (Paul Smith September 2007).

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Gallery 6 Heliopetes, Heliopyrgus and Melanion
FIGURE 6a - Alana´s White Skipper Heliopetes alana male - PROCOSARA, PN San Rafael (Paul Smith March 2007).
FIGURE 6b - Heliopetes arsalte - Encarnaciòn, Departamento Itapúa (Paul Smith August 2007).
FIGURE 6c - Heliopetes libra female - PROCOSARA, PN San Rafael (Paul Smith February 2008).
FIGURE 6d - Heliopetes omrina - PROCOSARA, PN San Rafael (Paul Smith January 2008).
FIGURE 6e - Heliopyrgus domicella - location unknown (Ulf Drechsel www.pybio.org).
FIGURE 6f - White Shield Skipper Melanion leucaspis - PROCOSARA, PN San Rafael (Moriz Stiefel April 2008 ).

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Gallery 9 Pyrgus, Pythonides and Quadrus
FIGURE 9a - Pluto´s Checkered Skipper Pyrgus orcus mating pair female upper, male lower - PROCOSARA, PN San Rafael (Paul Smith May 2008).
FIGURE 9b - Common Checkered Skipper Pyrgus orcynoides male - PROCOSARA, PN San Rafael (Paul Smith March 2007).
FIGURE 9c - Common Checkered Skipper Pyrgus orcynoides female - PROCOSARA, PN San Rafael (Martijn Brouwer May 2007).
FIGURE 9d - Blue Lance Skipper Pythonides lancea - PROCOSARA, PN San Rafael (David Gill February 2008).
FIGURE 9e - Ceres´ Skipper Quadrus cerialis - PROCOSARA, PN San Rafael (Sylvia Qu May 2008).
FIGURE 9f - Quadrus u-lucida - location unknown (Ulf Drechsel www.pybio.org).

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Gallery 12 Urbanus and Xenophanes
FIGURE 12a - Turquoise Long-tailed Skipper Urbanus proteus - upper photo Arroyo Mboi Kae, Departamento Itapúa (Regis Nossent July 2005), lower photo Encarnación, Departamento Itapúa (Paul Smith May 2007).
FIGURE 12b - Plain Long-tailed Skipper Urbanus simplicus - Encarnación, Departamento Itapúa (Paul Smith May 2007).
FIGURE 12c - Lined Long-tailed Skipper Urbanus teleus - upper photo Arroyo Mboi Kae, Departamento Itapúa (Regis Nossent July 2005), lower photo Encarnación, Departamento Itapúa (Paul Smith April 2009).
FIGURE 12d - Urbanus zagorus - PROCOSARA, PN San Rafael (Paul Smith November 2007).
FIGURE 12e - Glossy-winged Skipper Xenophanes tryxus - Encarnación, Departamento Itapúa (Paul Smith April 2009).

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Gallery 4 Cogia, Dyscophellus, Ebrietas, Epargyreus and Errynis
FIGURE 4a - Cogia calchas - Encarnación, Departamento Itapúa (Paul Smith January 2008).
FIGURE 4b - Cogia hassan - Estancia Laguna Blanca, Departamento San Pedro (Paul Smith December 2009).
FIGURE 4c - Dyscophellus erythras female - Hotel Tirol, Departamento Itapúa (Regis Nossent June 2005).
FIGURE 4d - Black-patched Duskywing Ebrietas anacreon - PROCOSARA, PN San Rafael (Paul Smith March 2007).
FIGURE 4e - Colossus Skipper Epargyreus clavicornis - PROCOSARA, PN San Rafael (Paul Smith January 2008).
FIGURE 4f - Errynis funeralis - Bahía de Asunción, Departamento Central (Paul Smith October 2009).

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Gallery 8 Pellicia, Phocides, Polythrix, Polygonus and Proteides
FIGURE 8a - Pellicia vecina - location unknown (Ulf Drechsel www.pybio.org).
FIGURE 8b - Black Boatman Phocides charon - PROCOSARA, PN San Rafael (Paul Smith September 2008).
FIGURE 8c - Red-shouldered Boatman Phocides polybus - location unknown (Sigrid Drechsel www.pybio.org).
FIGURE 8d - Polygonus leo - Estancia Kanguery, PN San Rafael (Paul Smith November 2007).
FIGURE 8e - Polygonus savigny - Itabó Itaipú Reserve, Departamento Alto Paraná (Paul Smith November 2008).
FIGURE 8f - Polythrix caunus - location unknown (Sigrid Drechsel www.pybio.org).
FIGURE 8g - Polythrix octomaculata - Estancia Laguna Blanca, Departamento San Pedro (Paul Smith February 2010).
FIGURE 8h - Mercurial Skipper Proteides mercurius - Estancia Laguna Blanca, Departamento San Pedro (Paul Smith February 2010).

Thanks to Ezequiel Nuñez-Bustos for assistance with butterfly identification.
Designed by Paul Smith 2006. This website is copyrighted by law.
Material contained herewith may not be used without the prior written permission of FAUNA Paraguay.
Photographs on this page were taken by Paul Smith, Martijn Brouwer,Regis Nossent, Laura Tensen, Jeni Oborn, Moriz Stiefel, Nick Simms, Sylvia Qu, David Gill, Ulf Drechsel, Sigrid Drechsel and Alberto Esquivel and are used with permission.
Special thanks to Ulf Drechsel for permission to use images from his site www.pybio.org.